Domestication History of Rye

Domestication History of Rye Rye (Secale cereale subspecies cereale) was likely fully domesticated from its weedy relative (S. cereale ssp segetale) or perhaps S. vavilovii, in Anatolia or the Euphrates River valley of what is today Syria, at least as early as 6600 BC, and perhaps as early as 10,000 years ago. Evidence for domestication is at Natufian sites such as Can Hasan III in Turkey at 6600 cal BC (calendar years BC); domesticated rye reached central Europe (Poland and Romania) about 4,500 cal BC. Today rye is grown on about 6 million hectares in Europe where it is mostly used for making bread, as animal feed and forage, and in the production of rye and vodka. Prehistorically rye was used for food in a variety of ways, as animal fodder and for straw for the thatched rooves. Characteristics Rye is a member of the Triticeae tribe of Pooideae subfamily of the Poaceae grasses, meaning it is closely related to wheat and barley. There are around 14 different species of the Secale genus, but only S. cereale is domesticated. Rye is allogamous: its reproductive strategies promote outcrossing. Compared to wheat and barley, rye is relatively tolerant to frost, drought, and marginal soil fertility. It has an enormous genome size (~8,100 Mb), and its resistance to frost stress appears to be a result of the high genetic diversity among and within rye populations. The domestic forms of rye have larger seeds than wild forms as well as a non-shattering rachis (the part of the stem that holds the seeds onto the plant). Wild rye is free-threshing, with a tough rachis and loose chaff: a farmer can free the grains by a single threshing since straw and chaff are eliminated by a single round of winnowing. Domestic rye maintained the free-threshing characteristic and both forms of rye are vulnerable to ergot and to munching by pesky rodents while still ripening. Experimenting with Rye Cultivation There is some evidence that Pre-Pottery Neolithic (or Epi-Paleolithic) hunters and gatherers living in the Euphrates valley of northern Syria cultivated wild rye during the cool, arid centuries of the Younger Dryas, some 11,000-12,000 years ago. Several sites in northern Syria show that increased levels of rye were present during the Younger Dryas, implying that the plant must have been specifically cultivated to survive. Evidence discovered at Abu Hureyra (~10,000 cal BC), TellAbr (9500-9200 cal BC), Mureybet 3 (also spelled Murehibit, 9500-9200 cal BC), Jerf el Ahmar (9500-9000 cal BC), and Djade (9000-8300 cal BC) includes the presence of multiple querns (grain mortars) placed in food processing stations and charred wild rye, barley, and einkorn wheat grains. In several of these sites, rye was the dominant grain. Ryes advantages over wheat and barley are its ease of threshing in the wild stage; it is less glassy than wheat and can be more easily prepared as food (roasting, grinding, boiling and mashing). Rye starch is hydrolyzed to sugars more slowly and it produces a lower insulin response than wheat, and is, therefore, more sustaining than wheat. Weediness Recently, scholars have discovered that rye, more than other domesticated crops has followed a weedy species type of domestication processfrom wild to weed to crop and then back to weed again. Weedy rye (S. cereale ssp segetale) is distinctive from the crop form in that it includes stem shattering, smaller seeds and a delay in flowering time. It has been found to have spontaneously redeveloped itself out of the domesticated version in California, in as few as 60 generations. Sources This article is part of the About.com guide to Plant Domestication, and part of the Dictionary of Archaeology Hillman G, Hedges R, Moore A, Colledge S, and Pettitt P. 2001. New evidence of Late Glacial cereal cultivation at Abu Hureyra on the Euphrates. The Holocene 11(4):383-393. Li Y, Haseneyer G, Schà ¶n C-C, Ankerst D, Korzun V, Wilde P, and Bauer E. 2011. High levels of nucleotide diversity and fast decline of linkage disequilibrium in rye (Secale cerealeL.) genes involved in frost response. BMC Plant Biology 11(1):1-14. http://dx.doi.org/10.1186/1471-2229-11-6 (Springer link is currently not working) Marques A, Banaei-Moghaddam AM, Klemme S, Blattner FR, Niwa K, Guerra M, and Houben A. 2013. B chromosomes of rye are highly conserved and accompanied the development of early agriculture. Annals of Botany 112(3):527-534. Martis MM, Zhou R, Haseneyer G, Schmutzer T, Vrna J, Kubalkov M, Kà ¶nig S, Kugler KG, Scholz U, Hackauf B et al. 2013. Reticulate Evolution of the Rye Genome. The Plant Cell 25:3685-3698. Salamini F, Ozkan H, Brandolini A, Schafer-Pregl R, and Martin W. 2002. Genetics and geography of wild cereal domestication in the near east. Nature Reviews Genetics 3(6):429-441.   Shang H-Y, Wei Y-M, Wang X-R, and Zheng Y-L. 2006. Genetic diversity and phylogenetic relationships in the rye genus Secale L. (rye) based on Secale cereale microsatellite markers. Genetics and Molecular Biology 29:685-691. Tsartsidou G, Lev-Yadun S, Efstratiou N, and Weiner S. 2008. Ethnoarchaeological study of phytolith assemblages from an agro-pastoral village in Northern Greece (Sarakini): development and application of a Phytolith Difference Index. Journal of Archaeological Science 35(3):600-613. Vigueira CC, Olsen KM, and Caicedo AL. 2013. The red queen in the corn: agricultural weeds as models of rapid adaptive evolution. Heredity 110(4):303-311.   Willcox G. 2005. The distribution, natural habitats, and availability of wild cereals in relation to their domestication in the Near East: multiple events, multiple centres. Vegetation History and Archaeobotany 14(4):534-541. http://dx.doi.org/10.1007/s00334-005-0075-x (Springer link not working) Willcox G, and Stordeur D. 2012. Large-scale cereal processing before domestication during the 10th millennium Cal BC in northern Syria. Antiquity 86(331):99-114.

A Summary of British Rule in India

A Summary of British Rule in India The very idea of the British Raj- the British rule over India- seems inexplicable today. Consider the fact that Indian written history stretches back almost 4,000 years, to the civilization centers of the Indus Valley Culture at Harappa and Mohenjo-Daro. Also, by 1850, India had a population of at least 200 million. Britain, on the other hand, had no indigenous written language until the 9th century C.E. (almost 3,000 years after India). Its population was about 16.6 million in 1850. How, then, did Britain manage to control India from 1757 to 1947? The keys seem to have been superior weaponry, economic power, and Eurocentric confidence. European Scramble for Colonies in Asia After the Portuguese rounded the Cape of Good Hope on Africas southern tip in 1488, opening sea lanes to the Far East by piracy on ancient trade lines in the Indian Ocean, the European powers strove to acquire Asian trading posts of their own. For centuries, the Viennese had controlled the European branch of the Silk Road, reaping enormous profits from the sale of silk, spices, fine china, and precious metals. The Viennese monopoly ended with the establishment of European incursions in the sea trade. At first, the European powers in Asia were solely interested in trade, but over time they became more interested in acquiring territory. Among the nations looking for a piece of the action was Britain. The Battle of Plassey Britain had been trading in India since about 1600, but it did not begin to seize large sections of land until 1757, after the Battle of Plassey. This battle pitted 3,000 soldiers of the British East India Company against the 5,000-strong army of the young Nawab of Bengal, Siraj ud Daulah, and his French East India Company allies. Fighting began on the morning of June 23, 1757. Heavy rain spoiled the Nawabs cannon powder (the British covered theirs), leading to his defeat. The Nawab lost at least 500 troops, while Britain lost only 22. Britain seized the modern equivalent of about $5 million from the Bengali treasury and used it to finance further expansion. India Under the East India Company The East India Company was primarily interested in trade in cotton, silk, tea, and opium, but following the Battle of Plassey, it functioned as the military authority in growing sections of India as well. By 1770, heavy Company taxation and other policies had left millions of Bengalis impoverished. While British soldiers and traders made their fortunes, the Indians starved. Between 1770 and 1773, about 10 million people (one-third  of the population) died of famine in Bengal. At this time, Indians were also barred from holding high office in their own land. The British considered them inherently corrupt and untrustworthy. The Indian Mutiny of 1857 Many Indians were distressed by the rapid cultural changes imposed by the British. They worried that Hindu and Muslim India would be Christianized. In 1857, a new type of rifle cartridge was given to the soldiers of the British Indian Army. Rumors spread that the cartridges had been greased with pig and cow fat, an abomination to both major Indian religions. On May 10, 1857, the Indian Revolt began, with Bengali Muslim troops marching to Delhi and pledging their support to the Mughal emperor. After a year-long struggle, the rebels surrendered on June 20, 1858. Control of India Shifts to the India Office Following the rebellion, the British government abolished both the Mughal Dynasty, which had ruled India more or less for 300 years, and the East India Company. The Emperor, Bahadur Shah, was convicted of sedition and exiled to Burma. Control of India was given over to a British Governor-General, who reported back to the British Parliament. It should be noted that the British Raj included only about two-thirds of modern India, with the other portions under the control of local princes. However, Britain exerted great pressure on these princes, effectively controlling all of India. Autocratic Paternalism Queen Victoria promised that the British government would work to better its Indian subjects. To the British, this meant educating the Indians in British modes of thought and stamping out cultural practices such as sati- the practice of immolating a widow on the death of her husband .The British thought of their rule as a form of autocratic paternalism. The British also created divide and rule policies, pitting Hindu and Muslim Indians against one another. In 1905, the colonial government divided Bengal into Hindu and Muslim sections; this division was revoked after strong protests. Britain also encouraged the formation of the Muslim League of India in 1907. British India During World War I During World War I, Britain declared war on Germany on Indias behalf, without consulting Indian leaders. More than 1.3 million Indian soldiers and laborers were serving in the British Indian Army by the time of the Armistice. A total of 43,000 Indian and Gurkha soldiers were killed. Although most of India rallied to the British flag, Bengal and Punjab were less easy to control. Many Indians were eager for independence, and they were led in their struggle by an Indian lawyer and political newcomer known as  Mohandas Gandhi (1869–1948) In April 1919, more than 5,000 unarmed protesters gathered at Amritsar, in Punjab. British troops fired on the crowd, killing an estimated 1,500 men, women, and children, even though the official death toll of the Amritsar Massacre as reported was 379. British India During World War II When World War II broke out, India once again contributed hugely to the British war effort. In addition to troops, the princely states donated substantial amounts of cash. By the end of the war, India had an incredible volunteer army of 2.5 million men. About 87,000 Indian soldiers died in combat. The Indian independence movement was very strong by this time, and British rule was widely resented. Some 30,000 Indian POWs were recruited by the Germans and the Japanese to fight against the Allies in exchange for their freedom. Most Indians, however, remained loyal. Indian troops fought in Burma, North Africa, Italy, and elsewhere. The Struggle for Indian Independence Even as World War II raged on, Gandhi and other members of the Indian National Congress (INC) demonstrated against British rule. The 1935 Government of India Act had provided for the establishment of provincial legislatures across the colony. The Act also created a federal government for the provinces and princely states and granted the right to vote to about 10% of Indias male population. These moves toward limited self-governance only made India more impatient for true self-rule. In 1942, Britain sent an envoy to India, led by the British Labour politician Stafford Cripps (1889–1952), offering future dominion status in return for help recruiting more soldiers. Cripps may have made a secret agreement with the Muslim League, allowing Muslims to opt out of a future Indian state. Mahatma Gandhi Laughing with Granddaughters. Bettmann / Getty Images Arrests of Gandhi and INC Leadership Gandhi and the INC did not trust the British envoy and demanded immediate independence in return for their cooperation. When the talks broke down, the INC launched the Quit India movement, calling for the immediate withdrawal of Britain  from India. In response, the British arrested the INCs leadership, including Gandhi and his wife. Mass demonstrations were carried out across the country but were crushed by the British Army. Britain may not have realized it, but it was now just a matter of time before the British Raj came to an end. The soldiers who had joined Japan and Germany in fighting the British were put on trial at Delhis Red Fort in early 1946. A series of ten courts-martial were held, trying 45 prisoners on charges of treason, murder, and torture. The men were convicted, but huge public protests forced the commutation of their sentences. Hindu/Muslim Riots and Partition On August 17, 1946, violent fighting broke out between Hindus and Muslims in Calcutta. The trouble quickly spread across India. Meanwhile, cash-strapped Britain announced its decision to withdraw from India by June 1948. Sectarian violence flared again as independence approached. In June 1947, representatives of the Hindus, Muslims, and Sikhs agreed to divide India along sectarian lines. Hindu and Sikh areas remained part of India, while predominantly Muslim areas in the north became the nation of Pakistan. This division of territory was known as the Partition. Millions of refugees flooded across the border in each direction, and between 250,000 and 500,000 people were killed in sectarian violence. Pakistan became independent on August 14, 1947. India followed the next day. Sources and Further Reading Gilmour, David. The British in India: A Social History of the Raj. New York: Farrar, Straus and Giroux, 2018.  James, Lawrence. Raj: The Making and Unmaking of British India. New York: St. Martins Griffin, 1997.Nanda, Bal Ram. Gokhale: The Indian Moderates and the British Raj. Princeton NJ: Princeton University Press, 1977.  Ã‚  Tharoor, Shashi. Inglorious Empire: What the British Did to India. London: Penguin Books Ltd, 2018.